| Taxa: |
| Order: |
| Family: |
| Mammalia |
| Chiroptera |
| Vespertilionidae |
| NatureServe Global Rank: |
| NatureServe State (NC) Rank: |
| G5 |
| S5B,SZN |
| Federal Status: |
| NC State Status: |
| --- |
| --- |
| Land Unit |
| US Fish & Wildlife Service |
| US Forest Service |
| US National Park Service |
| US Department of Defense |
| NC State Parks |
| NC University System |
| NC Wildlife Res. Com. |
| NC Forest Service |
| NC Div. of Coastal Mgmt. |
| Local Governments |
| Non-Governmental Org. |
| Other Public Lands |
| Private Lands |
| GAP Status 1-2 |
| All Protected Lands |
| Statewide |
| Hectares |
| 130,240.26 |
| 475,716.69 |
| 154,401.66 |
| 130,775.58 |
| 51,974.64 |
| 37,652.04 |
| 136,996.20 |
| 18,633.57 |
| 8,550.90 |
| 12,755.52 |
| 30,602.70 |
| 6,344.46 |
| 11,036,719.44 |
| 494,774.88 |
| 1,188,134.70 |
| 12,231,363.66 |
| Acres |
| 321,830.63 |
| 1,175,521.31 |
| 381,534.74 |
| 323,153.43 |
| 128,432.11 |
| 93,040.20 |
| 338,524.92 |
| 49,603.16 |
| 21,129.73 |
| 31,519.57 |
| 75,620.90 |
| 15,677.50 |
| 27,272,322.36 |
| 1,226,173.73 |
| 2,939,502.82 |
| 30,227,910.56 |
| % of Dist. on |
| Prot. Lands |
| 11.0 % |
| 40.0 % |
| 12.8 % |
| 11.0 % |
| 4.4 % |
| 3.2 % |
| 11.4 % |
| 1.6 % |
| 0.7 % |
| 2.5 % |
| 2.5 % |
| 0.4 % |
| < 0.1 % |
| 41.6 % |
| ----- |
| ----- |
| % of Dist. on |
| All Lands |
| 1.1 % |
| 3.9 % |
| 1.3 % |
| 1.1 % |
| 0.4 % |
| 0.3 % |
| 1.1 % |
| 0.2 % |
| < 0.1 % |
| 0.1 % |
| 0.3 % |
| < 0.1 % |
| 90.2 % |
| 4.0 % |
| ----- |
| ----- |
|
Ancestrally, roosts of N. humeralis are presumed to have been predominantly in hollow trees. However, the species has adapted well to manmade structures and now uses both types of shelter, arboreal and manmade, as roosts and nurseries (Whitaker and Hamilton 1998). Roosts and nurseries have been found in hollow trees, old and abandoned buildings,
house attics, barns, bridges, water culverts and other places that provide dark, secluded 'nooks and crannies' (Webster et al. 1985, Brown 1997). NATURE SERVE GLOBAL HABITAT COMMENTS: Prefers deciduous and mixed forest interspersed with cultivated areas. Commonly found along waterways (Schmidly 1991). Summer roosts include attics of buildings, tree cavities, and spaces behind loose tree bark. May also use Spanish moss for summer roosting, and cave mouths in fall. Hibernation site not known. Maternity colonies may be found under loose bark, in tree cavities, in Spanish-moss, or in buildings. |
| Code | Name | Description | NC Natural Heritage Program Equivalent |
| 75 | Tidal Swamp Forest | Swamp tupelo dominated forest with or without black tupelo and/or cypress trees. Restricted to the tidal zones in the coastal plain. May have inclusions of coastal red cedar woodlands. | Tidal cypress - gum swamp |
| 121 | Maritime Pinelands | Loblolly forests and woodlands of the outer coastal plain. | Estuarine Fringe Loblolly Pine Forest |
| 17 | Maritime Forests and Hammocks | Maritime forests and woodlands dominated by live or sand laurel oak. Estuarine Fringe forests dominated by loblolly pine. | Coastal Fringe Evergreen Forest, Maritime Deciduous Forest, Maritime Deciduous Forest |
| 126 | Interdune Wooded Depression Swamp | Includes swamps dominated by sweetbay and swampbay or dogwood dominated forests. | Maritime Shrub Swamp, Maritime Swamp Forest |
| 380 | Coastal Plain Fresh Water Emergent | Emergent vegetation in fresh water seepage bogs, ponds and riverbeds of the coastal plain. Includes alliances dominated by sedges, eelgrass, as well as cane found in unforested cane-brakes. | Small Depression Pond, Sandhill Seep, Floodplain Pool, Unforested Floodplain Canebrake, Riverscour Prairies, Vernal Pools |
| 173 | Coastal Plain Riverbank Shrubs | Shrub dominated riverbanks, commonly dominated by willows and/or alders. | Sand and Mud Bar |
| 50 | Coastal Plain Mixed Bottomland Forests | Includes forests dominated by a variety of hardwood species, including sweetgum, cottonwood, red maple. | Coastal Plain Bottomland Hardwood (in part), Coastal Plain Levee Forest |
| 49 | Coastal Plain Oak Bottomland Forest | Bottomland forests dominated by deciduous oak alliances. Oaks represented can include swamp chestnut, cherrybark, willow, and/or overcup oak. Inclusions of loblolly pine temporarily flooded forests occur in patches. Hydrology is temporarily to seasonally flooded. | Coastal Plain Bottomland Hardwoods (in part) blackwater subtype, brownwater subtype |
| 158 | Coastal Plain Nonriverine Wet Flat Forests | Loblolly pine - Atlantic white-cedar - red maple - swamp tupelo saturated forests as well as forests dominated by loblolly, sweetgum, and red maple in non-riverine flats. | Non-riverine Wet Hardwood Forest |
| 41 | Peatland Atlantic White-Cedar Forest | Dense stands of Atlantic white cedar with saturated hydrology. Can include swamp tupelo, red maple, and pond pines with a moderate shrub and herb layer. | Peatland Atlantic White-Cedar Forest |
| 15 | Seepage and Streamhead Swamps | Includes extensive peat flats in the coastal plain, dominated by swamp tupelo, maples, and Atlantic white cedar alliances. In the sandhills includes streamhead pond pine and bay forests alliances. Saturated hydrology. | Bay Forest, Small Depression Pocosin, Streamhead Atlantic White Cedar Forest, Streamhead Pocosins |
| 30 | Cypress-Gum Floodplain Forests | Swamps dominated by black or swamp tupelo with or without Taxodium. Seasonally to semi-permanently flooded hydrology. | Cypress-Gum Swamps |
| 78 | Pond-Cypress - Gum Swamps, Savannas and Lakeshores | Cypress dominated swamps and lakeshores. Can include bays dominated by pond cypress or shorelines of coastal plain lakes with a narrow band of cypress. | Non-riverine Swamp Forest, Natural Lakeshores (in part) |
| 385 | Oak Bottomland Forest and Swamp Forest | The swamp chestnut oak, cherrybark oak, shumard oak and sweetgum alliance is one representative. Other alliances are dominated by water, willow, and overcup oaks. Swamp forests can be dominated by sweetgum, red maple, and black gum being dominant. Loblolly can occur in combination with sweetgum and red maple, or with tulip poplar. Includes saturated and semi- to permanently flooded forests in the mountains. | Piedmont/Mountain Bottomland Forest, Piedmont/Mountain Swamp Forest |
| 63 | Coastal Plain Mesic Hardwood Forests | Beech dominated forests with white oak and northern red oak as possible co-dominants. Dry-mesic to mesic forests on slopes and small stream bottoms in the coastal plain. | Mesic Mixed Hardwood Forest, Basic Mesic Forests |
| 138 | Coastal Plain Dry to Dry-Mesic Oak Forests | Oak dominated forests of the coastal plain. Includes white oak forests with water oak or northern red oak and hickories as co-dominants. | Dry Mesic Oak Hickory Forest, Basic Oak Hickory Forest, Dry Oak Hickory Forest |
| 87 | Pocosin Woodlands and Shrublands | Includes pond pine woodland, low pocosin and high pocosin shrub dominated areas. Canebrakes and bay forests may be present. | Pond Pine Woodlands, Peatland Canebrake, Small Depression Pocosin |
| 67 | Wet Longleaf or Slash Pine Savanna | Wet flatwoods and pine savannas, typically dominated by longleaf pines, but slash or pond pines may be the dominant pines. | Wet Pine Flatwoods |
| 97 | Mesic Longleaf Pine | Longleaf pine woodlands without a major scrub oak component. Slash or loblolly pines may be present as well. | Mesic Pine Flatwoods |
| 42 | Xeric Longleaf Pine | Sandhills including a range of longleaf pine density from predominantly wiregrass, scrub oak dominated to true longleaf pine woodland. This does not include mesic or saturated flatwood types. | Xeric Sandhill Scrub, Pine/Scrub Oak Sandhill, Coastal Fringe Sandhill |
| 46 | Xeric Oak - Pine Forests | Mixed forest dominated by yellow pines with white or northern red oaks co-dominating. | Pine Oak Heath |
| 232 | Xeric Pine-Hardwood Woodlands and Forests | Mixed forest dominated by yellow pines with drier oaks including southern red, post, and chestnut oaks. | Dry Oak Hickory Forest |
| 238 | Piedmont/Mountain Submerged Aquatic Vegetation | Seasonally to permanently flooded areas with aquatic vegetation. Waterlily, pondweed, hydrilla smartweed are a few of the species that can occur. | Piedmont/Mountain Semipermanent Impoundment (in part) |
| 239 | Piedmont/Mountain Emergent Vegetation | Emergent vegetation of all wetland hydrologies. Sites would commonly support species such as tussock sedge, rushs, and cattail alliances. | Rocky Bar and Shore (in part) |
| 267 | Riverbank Shrublands | Riverside shrubs with temporarily flooded hydrologies. Found in the both the Mountains and Piedmont. Containing dominants such as smooth alder and a Carolina or black willows. | Sand and Mud Bar |
| 269 | Floodplain Wet Shrublands | Saturated shrublands of the Piedmont, includes buttonbush, swamp-loosestrife, decodon and alders. | Piedmont/mountain Semipermanent Impoundment |
| 230 | Piedmont Mesic Forest | American Beech - Red Oak - White Oak Forests. | Mesic Mixed Hardwood |
| 384 | Piedmont/Mountain Mixed Bottomland Hardwood Forests | Includes temporarily to seasonally forests dominated by hardwood species. Hardwoods include sweetgum, red maple, sycamore which co-occur in a mosaic of bottomland and levee positions. Includes alluvial hardwood forests in the mountains. Hemlock and white pine may occur as inclusions, but are generally mapped separately. | Piedmont/Mountain Alluvial Forest, Piedmont/Mountain Levee Forest |
| 383 | Piedmont Mixed Successional Forest | Generally loblolly mixed with successional hardwoods. Sweetgum, tulip poplar and red maple are common co-dominants in these successional forests. | No equivalent |
| 228 | Piedmont Dry-Mesic Oak and Hardwood Forests | Primarily oak dominated forests, white oak is often dominant, with co-dominants including . Also represented by sweetgum and tulip poplar dominated forests. | Dry Mesic Oak Hickory Forest, Basic Oak Hickory Forest, Dry Oak Hickory Forest |
| 222 | Piedmont Dry-Mesic Pine Forests | Loblolly dominated forests resulting from succession following clearing. This type occurs on all moisture regimes following disturbance with the exception of the extremely xeric sites. | No equivalent |
| 382 | Dry Mesic Oak Pine Forests | Mixed forests of the coastal plain and piedmont. Includes loblolly pine with white, southern red and/or post oak and loblolly with water oak. On basic sites of the piedmont, eastern red cedar may co-occur with post, black, and blackjack oaks. | Dry Mesic Oak Hickory Forest, Xeric Hard Pan Forest, Chestnut Oak Forest, Dry Mesic Oak Hickory Forest, Dry Oak Hickory Forest |
| 220 | Piedmont Xeric Pine Forests | Dry to xeric pine forests dominated by Virginia pine, shortleaf pine or Eastern Red Cedar. | Pine Oak Heath |
| 226 | Piedmont Xeric Woodlands | Generally post and blackjack oak dominated woodlands. White ash and pignut hickory can be found in combination with Eastern red cedar on glades. | Xeric Hardpan Forest |
| 20 | Coniferous Regeneration | Regenerating pine stands. Predominantly loblolly pine, but slash and longleaf stands occur as well. | No equivalent |
| 21 | Coniferous Cultivated Plantation (natural / planted) | Managed pine plantations, densely planted. Most planted stands are loblolly, but slash and longleaf occur as well. | No equivalent |
| 51 | Deciduous Cultivated Plantation | Planted deciduous trees. Includes sweetgum and sycamore plantations. | No equivalent |
| 36 | Successional Deciduous Forests | Regenerating deciduous trees with a shrub stature. Commonly dominated by sweetgum, tulip poplars and maples. | No equivalent |
| 180 | Agricultural Crop Fields | Farm fields used for row crops. | No equivalent |
| 205 | Agricultural Pasture/Hay and Natural Herbaceous | Farm fields used for pasture grass or hay production, as well as old fields dominated by native and exotic grasses. | No equivalent |
| 202 | Residential Urban | Includes vegetation interspersed in residential areas. Includes lawns, mixed species woodlots, and horticultural shrubs. Vegetation accounts for between 20 - 70% of the cover. | No equivalent |
| 203 | Urban Low-Intensity Developed | Highly developed areas with vegetation accounting for < 20% of the cover. | No equivalent |
| 204 | Urban High-Intensity Developed and Transportation Corridors | Highly developed areas including infrastructure such as roads, railroads. Vegetation represents < 20% of the cover. | No equivalent |
| 8 | Open water | Open water without aquatic vegetation. | No equivalent |
| 517 | Hemlock Floodplain Forest | Alluvial forest with hemlock and/or white pine in mountains and western piedmont. Hydrology is generally temporarily to seasonally flooded. | Canada Hemlock Forest |
| 521 | Spruce/Fir Forest | High Elevation Frazer-Fir - Red Spruce, Red Spruce and Red-Spruce-Yellow Birch Forests. Tree densities included here include both woodland to forest density. Highly intermixed with Northern Hardwoods, Grassy Balds, and Shrub Balds. | Red Spruce--Fraser Fir Forest, Fraser Fir Forest |
| 522 | Northern Hardwoods | High Elevation forests including yellow birch, American beech, and yellow buckeye. Includes forests with Hemlock and Yellow Birch. | Northern Hardwoods Forest, Boulderfield Forest |
| 523 | Grassy Bald | High Elevation grassy balds including Pennsylvania sedge, mountain oatgrass, as well as shrubby areas dominated by Alleghany and smooth blackberry. | Grassy Bald |
| 524 | Shrub Bald | Variable phenologies, predominantly evergreen balds with rhododendon and Mountain laurels. Deciduous shrubs including green alder and Alleghany and smooth blackberry are included as well. Red Oak - Chestnut Oak Woodlands may be included in cases where the density of the woodland species is low and the shrub component is dense. | Heath Bald |
| 525 | Appalachian Oak Forest | A variety of oak forest types including Black, White, Scarlet Oaks in dry to mesic situations. Includes forests historically co-dominated by American Chestnut. | High Elevation Red Oak Forest, Montane White Oak Forest |
| 526 | Appalachian Cove Forest | Mixed Mesophytic forests of the mountains. Includes tuliptree, basswood, yellow buckeye and surgar maple. This class is mapped to include cove forests dominated or co-dominated by hemlock. | Rich Cove Forest, Acidic Cove Forest |
| 527 | Appalachian Hemlock | Upland hemlock forests of the moutains region. Vary from side slopes to steep slope positions. | Canada Hemlock Forest |
| 528 | Appalachian Xeric Pine Forest | Pine forests and woodlands on xeric sites. A variety of pines, including Virginia, Shortleaf, Eastern White Pine, Table Mountain and Pitch pine. Often small areas of dense pine within a matrix of Xeric Oak-Pine Forests. | Pine Oak Heath |
| 529 | Appalachian Xeric Mixed Forest | Mixed forests with Virginia, Shortleaf, Eastern White Pine, Table Mountain and Pitch pines in combination with xeric oak species. Oaks include, white, Southern Red, black, and rock chestnut. | Pine Oak Heath |
| 530 | Appalachian Xeric Deciduous Forest | Deciduous forests in the mountains dominated by Xeric Oak species. Species include, white, Southern red, black, and rock chestnut. | High Elevation Red Oak Forest, Montane White Oak Forest |
| 533 | Appalachian Swamp Forest | Evergreen and deciduous forests with saturated hydrologies. This class may contain a variety of trees species, including hemlock - red maple, pitch pine, and white pine forests. | Swamp Forest-Bog Complex, Southern Appalachian Bog, Southern Appalachian Fen |
| 534 | Appalachian Wet Shrubland/ Herbaceous | Saturated shrubs and herbaceous vegetation. Often mapped as an inclusion in Appalachian Swamp Forest. | Southern Appalachian Bog, Southern Appalachian Fen |
|
Schmidly, D. J. 1991. The bats of Texas. Texas A & M Univ. Press, College Station. 188 pp.
Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal Species of the World:a Taxonomic and Geographic Reference. Second Edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Brown, L. N. 1997. A guide to the mammals of the southeastern United States. University of Tennessee Press, Knoxville. xiv + 236 pp. Whitaker, J.O. Jr. and W.J. Hamilton, Jr. 1998. Mammals of the eastern United States. Cornell Univ. Press, Ithaca, New York. 583 pp. Lowery, G. H., Jr. 1974. The mammals of Louisiana and its adjacent waters. Louisiana State University Press, Baton Rouge. 565 pp. Barbour, R. W., and W. H. Davis. 1969. Bats of America. The University of Kentucky Press, Lexington, Kentucky. Watkins, L. C. 1972. NYCTICEIUS HUMERALIS. Am. Soc. Mamm., Mammalian Species No. 23:1-4. Schwartz, Charles W., and Elizabeth R. Schwartz. 1981. The wild mammals of Missouri. University of Missouri Press, Columbia. 356 pp. Baker, Rollin H. 1983. Michigan mammals. Michigan State University Press. 642 pp. Webster, W. D., J. F. Parnell and W. C. Biggs Jr. 1985. Mammals of the Carolinas, Virginia, and Maryland. The University of North Carolina Press, Chapel Hill, NC. Caire, W., J. D. Tyler, B. P. Glass, and M. A. Mares. Z. Marsh (illustrator). 1989. Mammals of Oklahoma. University of Oklahoma Press, Norman. Oklahoma. 567 pp. |
For more information please contact them at:
NC-GAP Analysis Project
Dept. of Zoology, NCSU
Campus Box 7617
Raleigh, NC 27695-7617
(919) 513-2853
www.basic.ncsu.edu/ncgap